Trinkaus, E. Early modern humans. Annu. Rev. Anthropol. 34, 207–230 (2005).
Petraglia, M. D., Haslam, M., Fuller, D. Q., Boivin, N. & Clarkson, C. Out of Africa: new hypotheses and evidence for the dispersal of Homo sapiens along the Indian Ocean rim. Ann. Hum. Biol. 37, 288–311 (2010).
Liu, W. et al. The earliest unequivocally modern humans in southern China. Nature 526, 696–699 (2015).
Li, Z. Y. et al. Late Pleistocene archaic human crania from Xuchang, China. Science 355, 969–972 (2017).
Martinón-Torres, M. et al. Homo sapiens in the Eastern Asian Late Pleistocene. Curr. Anthropol. 58, S000 (2017).
Zhang, D. J. et al. Denisovan DNA in Late Pleistocene sediments from Baishiya Karst Cave on the Tibetan Plateau. Science 370, 584–587 (2020).
Ni, X. J. et al. Massive cranium from Harbin in northeastern China establishes a new Middle Pleistocene human lineage. Innovation 2, 100130 (2021).
Harvati, K. & Reyes-Centeno, H. Evolution of Homo in the Middle and Late Pleistocene. J. Hum. Evol. 173, 103279 (2022).
Bae, C. J. et al. Moving away from “the Muddle in the Middle” toward solving the Chibanian puzzle. Evol. Anthropol. 33, e22011 (2024).
Bae, C. J. & Wu, X. J. Making sense of eastern Asian Late Quaternary hominin variability. Nat. Commun. 15, 9479 (2024).
Fu, Q. M. et al. Denisovan mitochondrial DNA from dental calculus of the >146,000-year-old Harbin cranium. Cell 188, 3919–3926 (2025).
Fu, Q. M. et al. The proteome of the late Middle Pleistocene Harbin individual. Science 389, 704–707 (2025).
McBrearty, S. & Brooks, A. S. The revolution that wasn’t: a new interpretation of the origin of modern human behaviour. J. Hum. Evol. 39, 453–563 (2000).
Henshilwood, C. S. et al. A 100,000-year-old ochre-processing workshop at Blombos Cave, South Africa. Science 334, 219–222 (2011).
Wadley, L., Hodgskiss, T. & Grant, M. Implications for complex cognition from the hafting of tools with compound adhesives in the Middle Stone Age, South Africa. Proc. Natl. Acad. Sci. USA. 106, 9590–9594 (2009).
Hoffmann, D. L., Angelucci, D. E., Villaverde, V., Zapata, J. & Zilhão, J. Symbolic use of marine shells and mineral pigments by Iberian Neandertals 115,000 years ago. Sci. Adv. 4, eaar5255 (2018).
Zaidner, Y. et al. Middle Pleistocene Homo behaviour and culture at 140,000 to 120,000 years ago and interactions with Homo sapiens. Science 372, 1429–1433 (2021).
Scerri, E. M. L. & Will, M. The revolution that still isn’t: the origins of behavioural complexity in Homo sapiens. J. Hum. Evol. 179, 103358 (2023).
Schick, K. D. & Zhuan, D. Early Paleolithic of China and eastern Asia. Evol. Anthropol. 2, 22–35 (1993).
Gao, X. & Norton, C. J. A critique of the Chinese ‘Middle Palaeolithic’. Antiquity 76, 397–412 (2002).
Braun, D. R., Norton, C. J. & Harris, J. W. K. in Asian Paleoanthropology: From Africa to China and Beyond (eds Norton, C. J. & Braun, D. R.) 41–48 (Springer, Verlach, (2010).
Gao, X. Paleolithic cultures in China: uniqueness and divergence. Curr. Anthropol. 54, 358–370 (2013).
Li, F. Fact or fiction: the Middle Palaeolithic in China. Antiquity 88, 1303–1309 (2014).
Seong, C. & Bae, C. J. The eastern Asian ‘Middle Palaeolithic’ revisited: a view from Korea. Antiquity 90, 1151–1165 (2016).
Hu, Y. et al. Late Middle Pleistocene Levallois stone-tool technology in southwest China. Nature 565, 82–85 (2019).
Li, H., Li, Z. Y., Gao, X., Kuman, K. & Sumner, A. Technological behaviour of the early Late Pleistocene archaic humans at Lingjing (Xuchang, China). Archaeol. Anthropol. Sci. 11, 3477–3490 (2019).
Lin, N. R. et al. Lithic miniaturization and hafted tools in early Late Pleistocene Salawusu, North China. J. Archaeol. Sci. Rep. 48, 103831 (2023).
Ruan, Q. J. et al. Quina lithic technology indicates diverse Late Pleistocene human dynamics in East Asia. Proc. Natl. Acad. Sci. USA 122, e2418029122 (2025).
Murray, A. S. & Wintle, A. G. Luminescence dating of quartz using an improved single-aliquot regenerative-dose protocol. Radiat. Meas. 32, 57–73 (2000).
Wintle, G. & Murray, A. S. A review of quartz optically stimulated luminescence characteristics and their relevance in single-aliquot regeneration dating protocols. Radiat. Meas. 41, 369–391 (2006).
Wang, X. L., Lu, Y. C. & Wintle, A. G. Recuperated OSL dating of fine-grained quartz in Chinese loess. Quat. Geochronol. 1, 89–100 (2006).
Kang, S. G., Lu, Y. C. & Wang, X. L. Closely-spaced recuperated OSL dating of the last interglacial paleosol in the southeastern margin of the Chinese Loess Plateau. Quat. Geochronol. 6, 480–490 (2011).
Buylaert, J.-P. et al. A robust feldspar luminescence dating method for Middle and Late Pleistocene sediments: Feldspar luminescence dating of Middle and Late Pleistocene sediments. Boreas 41, 435–451 (2012).
Buylaert, J.-P., Murray, A. S., Thomsen, K. J. & Jain, M. Testing the potential of an elevated temperature IRSL signal from K-feldspar. Radiat. Meas. 44, 560–565 (2009).
Keeley, L. H. Hafting and retooling: effects on the archaeological record. Am. Antiquity 47, 798–809 (1982).
Rots, V. in Multidisciplinary Approaches to the Study of Stone Age Weaponry (eds Iovita, R. & Sano, K.) 167–185 (Springer, 2016).
Tomasso, S. & Rots, V. What is the use of shaping a tang? Tool use and hafting of tanged tools in the Aterian of northern Africa. Archaeol. Anthropol. Sci. 10, 1389–1417 (2018).
Delpiano, D., Gravina, B. & Peresani, M. Back(s) to basics: the concept of backing in stone tool technologies for tracing hominins’ technical innovations. Evol. Anthropol. 33, e22045 (2024).
Rots, V. Prehension and Hafting Traces on Flint Tools: A Methodology (Leuven University Press, 2010).
Spry, C., Kurpiel, R., Foley, E. & Penzo-Kajewski, P. Revisiting the “quartz problem” in lithic studies: a review and new, open-access, experimental dataset. Lithic Technol. 47, 171–181 (2022).
Ma, D. D. et al. Earliest prepared core technology in Eurasia from Nihewan (China): Implications for early human abilities and dispersals in East Asia. Proc. Natl. Acad. Sci. USA. 121, e2313123121 (2024).
Yang, S. X. et al. Technological innovations at the onset of the Mid-Pleistocene Climate Transition in high-latitude East Asia. Natl. Sci. Rev. 8, nwaa053 (2021).
Hou, Y. M. et al. Mid-Pleistocene Acheulean-like stone technology of the Bose Basin, South China. Science 287, 1622–1626 (2000).
Li, H. et al. The temporal-spatial evolution of handaxe technology in China: Recent progress and future directions. Sci. Bull. 69, 2161–2165 (2024).
Li, H. et al. The Middle Pleistocene handaxe site of Shuangshu in the Danjiangkou Reservoir Region, Central China. J. Archaeol. Sci. 52, 391–409 (2014).
Pei, S. W. et al. Middle Pleistocene hominin occupation in the Danjiangkou Reservoir Region, Central China: Studies of formation processes and stone technology of Maling 2A site. J. Archaeol. Sci. 53, 391–407 (2015).
Liu, K. et al. Stratigraphy and chronology of two newly discovered Early Pleistocene Palaeolithic sites in the Hanjiang River Valley, Central China. Palaeogeogr. Palaeoclimatol. Palaeoecol. 605, 111229 (2022).
Gao, X. A study of flaking technology at Zhoukoudian Locality 15. Acta Anthropol. Sin. 19, 199–215 (2000).
Gao, X. A study of stone tool typology and retouch technology at Zhoukoudian Locality 15. Acta Anthropol. Sin. 20, 1–18 (2001).
Ren, J. C., Li, F., Chen, F. Y., Olsen, J. W. & Gao, X. Lithic technology at Banjingzi Locality 1: implications for the Middle Paleolithic in northern China. J. Archaeol. Sci. Rep. 65, 105193 (2025).
Liu, Y., Hou, Y. M. & Bao, L. The lithic industry and its cultural significance of the Wulanmulun site in Ordos region. Acta Archaeol. Sin. 4, 423–440 (2022).
Chen, H. et al. Hafting wear on quartzite tools: an experimental case from the Wulanmulun Site, Inner Mongolia of North China. Quatern. Int. 427, 184–192 (2017).
Li, H. et al. Continuous technological and behavioural development of Late Pleistocene hominins in central South China: multidisciplinary analysis at Sandinggai. Quat. Sci. Rev. 298, 107850 (2022).
Hu, Y., Ruan, Q., Liu, J., Marwick, B. & Li, B. Luminescence chronology and lithic technology of Tianhuadong Cave, an early Upper Pleistocene Paleolithic site in southwest China. Quat. Res. 94, 1–16 (2020).
Doyon, L., Li, Z. Y., Li, H. & d’Errico, F. Discovery of circa 115,000-year-old bone retouchers at Lingjing, Henan, China. PLoS ONE 13, e0194318 (2018).
Doyon, L., Li, H., Li, Z., Wang, H. & Zhao, Q. Further evidence of organic soft hammer percussion and pressure retouch from Lingjing (Xuchang, Henan, China). Lithic Technol. 44, 100–117 (2019).
Liu, J. H. et al. 300,000-year-old wooden tools from Gantangqing, southwest China. Science 389, 78–83 (2025).
Li, Z. Y. et al. Engraved bones from the archaic hominin site of Lingjing, Henan Province. Antiquity 93, 886–900 (2019).
Wu, X. J. et al. Evolution of cranial capacity revisited: a view from the late Middle Pleistocene cranium from Xujiayao, China. J. Hum. Evol. 163, 103119 (2022).
Bae, C. J., Li, F., Cheng, L. L., Wang, W. & Hong, H. L. Hominin distribution and density patterns in Pleistocene China: climatic influences. Palaeogeogr. Palaeoclimatol. Palaeoecol. 512, 118–131 (2018).
Liu, W. et al. Late Middle Pleistocene hominin teeth from Panxian Dadong, South China. J. Hum. Evol. 64, 337–355 (2013).
Lisiecki, L. E. & Raymo, M. E. A Pliocene-Pleistocene stack of 57 globally distributed benthic δ18O records. Paleoceanography 20, PA1003 (2005).
Hao, Q. et al. Delayed build-up of Arctic ice sheets during 400,000-year minima in insolation variability. Nature 490, 393–396 (2012).
Beck, J. W. et al. A 550,000-year record of East Asian monsoon rainfall from 10Be in loess. Science 360, 877–881 (2018).
D’Errico, F. & Banks, W. E. Identifying mechanisms behind Middle Paleolithic and Middle Stone Age cultural trajectories. Curr. Anthropol. 54, S371–S387 (2013).
Inizan, L., Reduron-Ballinger, M., Roche, H. & Tixier, J.Technology and Terminology of Knapped Stone (translated by J. Féblot-Augustins) (Cercle de Recherches et d’Etudes Préhistoriques, 1999).
Pelegrin, J. in Stone Knapping: The Necessary Conditions for a Uniquely Hominin Behaviour (eds Roux, V. & Bril, B.) 23–33 (McDonald Institute for Archaeological Research, 2005).
Soressi, M. & Geneste, J.-M. The history and efficacy of the chaîne opératoire approach to lithic analysis: Studying techniques to reveal past societies in an evolutionary perspective. PaleoAnthropology 334, 350 (2011).
Scerri, E. L., Gravina, B., Blinkhorn, J. & Delagnes, A. Can lithic attribute analyses identify discrete reduction trajectories? A quantitative study using refitted lithic sets. J. Archaeol. Method Th. 23, 669–691 (2016).
Ollé, A. & Vergès, J. M. The use of sequential experiments and SEM in documenting stone tool microwear. J. Archaeol. Sci. 48, 60–72 (2014).
Fernández-Marchena, J. L. et al. Rainbow in the dark. The identification of diagnostic projectile impact features on rock crystal. J. Archaeol. Sci. Rep. 31, 102315 (2020).
Fernández-Marchena, J. L. & Ollé, A. Microscopic analysis of technical and functional traces as a method for the use-wear analysis of rock crystal tools. Quatern. Int. 424, 171–190 (2016).
Pedergnana, A. & Ollé, A. Monitoring and interpreting the use-wear formation processes on quartzite flakes through sequential experiments. Quatern. Int. 427, 35–65 (2017).
Borel, A., Ollé, A., Vergès, J. M. & Sala, R. Scanning electron and optical light microscopy: two 1208 complementary approaches for the understanding and interpretation of usewear and residues on stone tools. J. Archaeol. Sci. 48, 46–59 (2014).
Fernández-Marchena, J. L. et al. Traceological analysis of a singular artefact: the rock crystal point from O Achadizo (Boiro, A Coruña, Galicia). J. Lithic Stud. 3, 253–271 (2016).
Martín-Viveros, J. I. & Ollé, A. Use-wear and residue mapping on experimental chert tools. A multi-1213 scalar approach combining digital 3D, optical, and scanning electron microscopy. J. Archaeol. Sci. Rep. 30, 102236 (2020).
Ollé, A. et al. Microwear features on vein quartz, rock crystal and quartzite: a study combining optical light and scanning electron microscopy. Quatern. Int. 424, 154–170 (2016).
Vergès, J. M. & Ollé, A. Technical microwear and residues in identifying bipolar knapping on an anvil: experimental data. J. Archaeol. Sci. 38, 1016–1025 (2011).
Pedergnana, A., Asryan, L., Fernández-Marchena, J. L. & Ollé, A. Modern contaminants affecting microscopic residue analysis on stone tools: a word of caution. Micron 86, 1–21 (2016).
Taipale, N. & Rots, V. Breakage, scarring, scratches and explosions: understanding impact trace formation on quartz. Archaeol. Anthropol. Sci. 11, 3013–3039 (2019).
Knutsson, K. Patterns of tools use: Scanning electron microscopy of experimental quartz tools. In Aun, 10. (Societas Archaeologica Upsalensis, Uppsala, 1988).
Pedergnana, A., García-Antón, M. D. & Ollé, A. Structural study of two quartzite varieties from the Utrillas facies formation (Olmos de Atapuerca, Burgos, Spain): from a petrographic characterisation to a functional analysis design. Quatern. Int. 433, 163–178 (2017).
Bustos-Pérez, G. & Ollé, A. The quantification of surface abrasion on flint stone tools. Archaeometry 66, 247–265 (2024).
Galland, A., Clemente-Conte, I., Boisserie, J.-R. & Delagnes, A. How depositional environments 1233 impact the microwear preservation of quartz artefacts: insights from the Oldowan of the Shungura Formation (Ethiopia). https://doi.org/10.31233/OSF.IO/9K8P3 (2024).
Ryan, W. B. F. et al. Global Multi-Resolution Topography (GMRT) synthesis data set. Geochem. Geophys. Geosyst. 10, Q03014 (2009).